... nine adult male Orang-utans, collected by myself in Borneo, the skulls differed remarkably in size and proportions. The orbits varied in width and height, the cranial ridge was either single or double, either much or little developed, and the zygomatic aperture varied considerably in size. I noted particularly that these variations bore no necessary relation to each other, so that a large temporal muscle and zygomatic aperture might exist either with a large ... — Darwinism (1889) • Alfred Russel Wallace